Often times after the music has been recorded, they will speed it up ever so slightly ... which can result in the recording being somewhat between keys.
Also, You make loving fun was initally recorded with John and Chris and Mick ...Chris playing her part on a Fender Rhodes. Unlike today's keyboards (or guitars), these vintage keyboards would often fall out of tune and are a real pain to re-tune. Perhaps John and Lindsey tuned their guitars to be in tune with her Rhodes, which may have been slightly flat.
Or even this possibility... these old albums were recorded on Old machines, old analog tapes, etc. .. Often times the speed of the tape machine can vary, so what may have been recorded on C or Bb, when replayed on a different machine for re-mastering the speed of the tape may have been slower thus taking the song down a quarter-tone or so
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